Mark Howarth is the Professor of Protein Nanotechnology in Oxford University Department of Biochemistry, where he has been a principal investigator since 2007. In 2017 he was awarded the Royal Society of Chemistry Norman Heatley Award. He did postdoctoral studies at MIT with Alice Ting, where he developed monovalent streptavidin and single molecule probes for tracking neurotransmitter receptors. His doctoral work was with Tim Elliott at Southampton University investigating MHC class I-peptide quality control. His current work is on innovating ultra-stable protein interactions for applications in synthetic biology, enzyme stabilization and vaccines.